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Echo is one of 900 elephants that live in Kenya's Amboseli National Park, an elderly matriarch and the leader of her family. Cynthia Moss knows them all, and with the help of cameraman Martyn Colbeck, made a unique record of Echo and her family. This film, narrated by David Attenborough, chronicles four years of Echo's life.

Echo is pregnant, her grand-daughter Edwina breaks a leg and her calf Ely becomes afflicted with 'twisting disease'. Meanwhile, the bull elephants of Amboseli - the biggest in Africa - suffer a tragedy, which raises the question of whether Ely will survive to breed himself.