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Nine celebrities trace their family trees to explore the lives of their ancestors and uncover major themes in British social history.

Rupert Everett's varied acting career has involved starring in Hollywood films as diverse as My Best Friend's Wedding, Shrek 2 and The Importance of Being Earnest. Rupert has a rigidly conservative background and feels that much of his own life has been a reaction against it. Following the recent death of his father six months ago, he wants to investigate the paternal side of his family.

Rupert visits his mother Sara to find out more about his father Tony's early life. He discovers that Tony's father Cyril worked in Nigeria for the Colonial Service. Sara has a photograph of Cyril as a two-year-old and tells Rupert that the family believe he was brought up by two aunts in Hammersmith. Rupert wants to find out more about Cyril's life and what he was doing in Nigeria.

Rupert tracks down records which chart Cyril's career in the Service and discovers just how important a role he held in Nigeria. But what about Cyril's childhood? Rupert decides to search the online census records and unearths some very unexpected information. This is the start of a rollercoaster investigation into fathers and sons which Rupert describes as like an Ealing comedy, with its twists, turns and surprise relatives popping up.

Rupert reflects on the ripple effects of broken relationships across the generations. On social mobility, both up and down, broken rules, abandonment, rebellion, convention and secrecy.