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Spring has sprung, and Verne and his woodland friends awaken from their long winter's nap to discover that a large, green hedge has cropped up right through the middle of their once-natural habitat. Enter RJ, an opportunistic raccoon who explains that the world beyond the hedge is 'the gateway to the good life' where peculiar creatures called humans live to eat, rather than eat to live. Verne, suspicious of RJ, believes that there is more to him than meets the eye and attempts to keep his family safely on their side of the hedge. Achieving that will not prove to be an easy feat.

Featuring the voices of Thomas Haden Church (Dwayne), Allison Janney (Gladys), Omid Djalili (Tiger), Avril Lavigne (Heather), Catherine O'Hara (Penny), Eugene Levy (Lou), Bruce Willis (RJ), Garry Shandling (Verne), Steve Carell (Hammy), Wanda Sykes (Stella), William Shatner (Ozzie) and Nick Nolte (Vincent).